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Alan Cumming is Alexander Novak, a man from Scotland who returns to the land of his forefathers in order to track down a piece of film showing his grandparents as they are taken away by the Nazi occupiers during World War II. Sandrine Bonnaire is a woman working at the film archives, whom Novak ends up falling in love with, which causes problems when her boyfriend (Bruno Ganz) enters the fray.
The feature-film début of actor Alan Cumming, Prague was selected at the Cannes Film Festival and was one of the few films set in the titular city from this time.